Puzzle 20

"Why so serious?", asked this villain

South __, the first state to secede from the Union

President who Lincoln succeeded, James __

Greenwich __, or GMT

Reverse direction

Online pages, e.g. Facebook, Twitter

A man who rents out property

Throwing away

Microscopic organisms that can cause disease

Not eaten

Tin full of watercolours

Thousands of millions

Main ingredient in pomodoro sauce

Progressive deterioration of brain functions

TV programme with quiz questions and prizes